British-Indian geneticist and polymath who was one of the founders of population genetics and the modern evolutionary synthesis — combining Mendelian genetics with Darwinian evolution.
Population genetics: mathematical treatment of evolution by natural selection operating on Mendelian genes — one of the three founders (with Fisher and Wright) of the mathematical synthesis. The Causes of Evolution (1932): integrated Mendelian genetics with natural selection. Kin selection: first mathematical treatment of the genetics of altruism (before Hamilton formalized it). Famous remark about God and beetles.
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